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Microscopy Confocal microscopy was performed using a Leica TCS SP8 confocal microscope (Leica, Wetzlar, Germany) equipped with a photomultiplier tube, two HyD hybrid detectors, and lasers at 405 nm, 488 nm, 552 nm, and 638 nm

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Acknowledgements This work was supported by a grant from the National Research Foundation of Korea (No