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The pro-inflammatory state (M1) is caused by pathogens, and macrophages are an important way to fight off an infection

One of the primary causes are free radicals which cause oxidation (rusting) of cell membranes, DNA, and so on

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Studies indicate potential involvement with the FAK-paxillin pathway, which plays a role in cell adhesion and tissue repair

Aging and Longevity Research Age-related changes in L-Carnitine metabolism and mitochondrial function connect to fundamental aging research: Mitochondrial Aging Research: Investigation of age-related mitochondrial dysfunction, decreased oxidative capacity, increased ROS production, and altered mitochondrial dynamics