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The sTNF--VNAR-C4 complex was crystallized by the sitting drop vapor diffusion method, using a reservoir solution containing 0.1 M MES, pH 6, 0.1 M magnesium chloride, and 8% w/v polyethylene glycol 6000

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It was first isolated in human plasma in 1973, and subsequent decades of research have revealed a mechanism that goes considerably beyond simple collagen stimulation