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On the one hand, ROS, as a second messenger, can activate MAPK or NF-B pathway and mediate granulocyte macrophage colony-stimulating factor (GM-CSF) production (224), which can interact with RNAKL and M-CSF to promote osteoclast differentiation (225)

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Acknowledgements This research was funded by the National Natural Science Foundation of China (81971187, 82171401, 81971183, and 82101477), Shanghai Municipal Science and Technology Major Project (2018SHZDZX05) and Innovation Program of Shanghai Municipal Education Commission (2017-01-07-00-01-E00046)