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Introduction Dihydroflavonol 4-reductase (DFR, EC1.1.1.219), a key enzyme in the regulation of the carbon flux direction in the flavonoid biosynthetic pathway, catalyzes the reduction of the 4-keto group of dihydroflavonol to the corresponding leucoanthocyanidin, which is the first committed reaction leading to anthocyanin production ( DFR genes leads to an increase in anthocyanin content, and the altered phenotypes of transgenic plants reflect the physiological roles of anthocyanins in plants ( The phytohormone auxin is critical for plant growth and it influences many developmental processes ( O -rhamnoside-7- O -rhamnoside) has been demonstrated to act as an endogenous polar auxin transport inhibitor in Arabidopsis ( Flowering time is essential for plant reproduction and is an important trait in agriculture