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More specifically, many genes encoding pathogenesis-related proteins such as osmotin-like protein ( OLP ), pathogenesis-related protein 1 ( PR1 ), pathogenesis-related protein 10 ( PR10 ), pathogenesis-related protein STH-2 ( STH2 ) and pathogenesis-related genes transcriptional activator 5 ( PTI5 ) were up-regulated in the seedlings treated with LNT/SPc complex (Fig

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